Leonardo da Vinci

The Florentine master behind the Last Supper and the Mona Lisa — his documented life, the long dispersal of the notebooks, the anatomy studies, and an audit of the 'universal genius' legend.

Red-chalk drawing traditionally identified as a self-portrait of Leonardo da Vinci
Red-chalk drawing traditionally identified as a self-portrait of Leonardo da Vinci; Royal Library, Turin. · Leonardo da Vinci (traditional identification) · Public domain

Leonardo da Vinci (15 April 1452 – 2 May 1519), born at Anchiano near Vinci in the Republic of Florence to unmarried parents — his father Ser Piero a Florentine notary, his mother Caterina a young woman of debated identity — was apprenticed at about fifteen to Andrea del Verrocchio, painted the angel of the Baptism of Christ, and was admitted to Florence's painters' guild in 1472. From 1482 to 1500 he served the Sforza court in Milan as painter and engineer, where he painted the Last Supper (1495–98) — an experimental tempera-on-dry-wall work that began flaking within years and was coated with later retouches until a 20-year restoration completed in 1999. Back in Florence he began the Mona Lisa about 1503, kept it unfinished until his death, and worked on anatomy in two documented campaigns (c. 1490 and 1506–13), dissecting about thirty human corpses; the anatomical treatise was never published in his lifetime. He died at Cloux (Clos-Lucé) in France on 2 May 1519, leaving more than 10,000 notebook pages to Francesco Melzi; the sheets were later bound by Pompeo Leoni into the Codex Atlanticus (Ambrosiana, Milan) and the Windsor volumes (Royal Collection, c. 550 drawings), while the Codex Leicester toured the Uffizi in 2018–19. Only 17 of his surviving paintings can be definitely attributed to him, and the attribution of works such as Salvator Mundi — sold in 2017 for $450.3 million — remains contested.

Birth and family in Vinci (1452)

Leonardo da Vinci was born on 15 April 1452 at Anchiano, near Vinci in the Republic of Florence1. He died on 2 May 1519 at Cloux — now Clos-Lucé — in France1. His parents were unmarried when he was born: his father, Ser Piero da Vinci, was a Florentine notary and landlord1. His mother was possibly Caterina di Meo Lippi, a young peasant woman who may have been orphaned early1. Though a few scholars have posited that she was an enslaved woman from Asia1. He grew up on his father's family estate, treated there as a legitimate son, with an estimated 22 half-siblings1.

His schooling was the elementary kind usual for the day — reading, writing and arithmetic1. Latin came later and largely by his own effort, and higher mathematics only from about the age of thirty1. He was left-handed, so mirror writing came easily and naturally to him1. His script, unusual though it was, can be read clearly with a mirror — as his contemporaries testified — and should not be looked on as a secret handwriting1.

Verrocchio's workshop and the early works

When he was about fifteen, his father apprenticed him to the artist Andrea del Verrocchio1. In that renowned workshop he received a multifaceted training in painting and sculpture as well as the technical-mechanical arts1. He also worked in the next-door workshop of Antonio Pollaiuolo1.

His talent — keenness of observation and creative imagination — was already apparent in the angel he contributed to Verrocchio's Baptism of Christ (c. 1470–75)1. Vasari narrates that Verrocchio was helped by a young pupil, Leonardo, who painted the angel on the left with such skill that he upset the older master2. Current studies regard his interventions as more extensive, including the riparian landscape, the golden light and the figure of Christ2.

In 1472 he was accepted into the painters' guild of Florence, stayed in his teacher's workshop for five more years, then worked independently in Florence until 14811. The Uffizi's Annunciation, considered a youthful work painted in Verrocchio's studio, dates to about 147231. In the portrait of Ginevra de' Benci he broke with the customary profile pose to show the young woman in three-quarter view31. The unfinished St. Jerome shows his early interest in anatomy31. When he left for Milan in 1482 he had just received the Adoration of the Magi commission, and one for an altar painting in the St. Bernard Chapel that was never begun1.

First Milan: the Sforza court (1482–1500)

Leonardo moved to Milan in 1482 to work in the service of the city's duke — a surprising step for the thirty-year-old, who had just received the Adoration of the Magi commission1. He was listed in the register of the duke's household as pictor et ingeniarius ducalis, 'painter and engineer of the duke'1. He spent seventeen years in Milan, until Ludovico Sforza's fall from power in 14991. Court festivals, fortifications and hydraulics fell within his brief1.

His most ambitious project was the equestrian monument to Francesco Sforza, pursued over twelve interrupted years1. A clay model was shown in 14931. Preparations were made to cast the colossal figure, which was to stand 16 feet (5 metres) high1. But because war was imminent the metal, ready to be poured, went to cannon instead, and the project came to a halt1. The model did not survive1.

The earliest painting he completed in Milan is probably The Virgin of the Rocks, an altarpiece for the church of San Francesco Grande begun about 14831. The project led to ten years of litigation with the Confraternity of the Immaculate Conception, and to a second version of the work completed in 15081. He completed at least six paintings during the seventeen Milan years, among them the portraits now called Lady with an Ermine and La Belle Ferronnière1.

The Last Supper (1495–98): technique, damage, restoration

The Last Supper was painted by Leonardo probably between 1495 and 1498 for the refectory of the Dominican monastery Santa Maria delle Grazie in Milan4. Ludovico Sforza, his patron during the first Milanese stay, commissioned it4. The Sforza coats of arms appear with the family's initials on the lunettes above the mural4. The scene fixes the moment Jesus announces that one of the Apostles will betray him4. Jesus and Judas reach toward the same dish between them — an act that marks Judas as the betrayer4.

Leonardo bypassed true fresco, which requires painting quickly on fresh plaster, and tried instead an experimental technique of tempera or oil paint on two layers of dry preparatory ground4. The pigments were not permanently attached to the wall4. The painting began to flake within a few years4. The deterioration continued for centuries — the kitchen's steam and smoke, the refectory's candle soot, the dampness of the location4. A door cut through the wall in 1652 through the feet of Jesus, Napoleonic troops stabled horses in the refectory, and a nineteenth-century flood and mould followed4. During the Second World War an Allied bomb collapsed the roof and one wall, leaving the painting exposed to the elements for several months before the space was rebuilt4.

Date / periodEvent
1495–98Painted for the refectory of Santa Maria delle Grazie.4
Early 1500sThe paint begins to flake: the pigments never bonded to the wall.4
1652A door is cut through the wall, through the feet of Jesus.4
Napoleonic eraTroops stable horses in the refectory.4
Early 19th centuryFlood and mould cause further damage.4
World War IIAn Allied bomb collapses the roof and one wall; the painting is exposed for several months.4
1999A 20-year restoration is completed, amid fierce debate.4
TodaySmall groups of visitors are admitted for fifteen minutes.4
The Last Supper: painting, damage and conservation, 1495–1999.

After centuries of maltreatment the mural underwent an extensive and controversial twenty-year restoration completed in 19994. Restorers removed previous retouches, grime and varnish, and many critics argued that very little of Leonardo's original work was left4. The Cenacolo museum describes the masterpiece, shaped by its technique, its environment and its historical vicissitudes, as vulnerable and fragile5. Its staff care for it daily with a wide range of preventive actions5. Visitors are admitted in small groups, with fifteen minutes allotted for the viewing4.

The notebooks and their dispersal

Leonardo began the earliest of his notebooks during his first years in Milan, sketching his observations on loose sheets or on the tiny paper pads he kept in his belt1. At various points he tried to arrange the notes by theme1. Four themes were planned — painting, architecture, mechanics and anatomy1. Drawing takes precedence over text in the manner of a dimostrazione1. Throughout, the script runs in his left-handed mirror writing1.

Because Leonardo never got around to publishing his scientific and artistic studies, his loyal student and friend Melzi inherited his estate1. More than ten thousand closely written and abundantly illustrated pages, the most voluminous literary legacy any painter had ever left behind1. About twenty-one codices have survived, contemporary sources mention forty, and thirty-two have been preserved in all once the later bindings are counted1. After Melzi's death around 1570 the collection passed to the sculptor Pompeo Leoni6. He arranged the drawings loosely by subject and pasted them into the pages of at least two large albums6.

Manuscript / collectionLocationKey figures
Codex AtlanticusVeneranda Biblioteca Ambrosiana, Milan1,200 folios; collected by Pompeo Leoni at the end of the 16th century.7
Royal Library albumWindsor Castlec. 600 Vincian sheets; c. 550 Leonardo drawings in the Royal Collection.76
Madrid CodicesNational Library of MadridTwo notebooks discovered in 1965.1
Codex LeicesterExhibited at the Uffizi, 2018–1918 sheets folded into 72 pages; main parts 1504–1508.8
Principal Leonardo manuscripts and where they are documented today.

The Codex Atlanticus, the largest collection of Leonardo's writings and drawings, is preserved at the Veneranda Biblioteca Ambrosiana in Milan, catalogued at 1,200 folios97. The Royal Collection holds around 550 of his drawings — the most important group in the world — kept in boxes at Windsor because light can damage them6. The so-called Madrid Codices were discovered in 1965 in the National Library of Madrid1. The Codex Leicester, eighteen sheets folded into a 72-page document whose main parts originated between 1504 and 15088. It was shown at the Uffizi from 30 October 2018 until 20 January 20198. Melzi is believed to have compiled the Trattato della pittura before 1542110. It was first printed in 1651 and published complete only in 1817, while the anatomical drawings reached print in the late nineteenth century110. The Museo Galileo's leonardo//thek@ project put the Codex Atlanticus online in March 20237. Its 2.0 version, due in 2026, adds the 600 Vincian sheets at Windsor — together one-third of the extant corpus7.

Anatomy: dissections, drawings, and their dating

Leonardo's anatomical work belongs to two campaigns of research — around 1490, and between 1506 and 151310. In them he investigated the nervous system, the internal organs, the bones and muscles, the structure and function of the heart, and the reproductive systems10. What began as training for an artist had grown by the 1490s into an independent area of research1. The dissections themselves may have begun several years after he first moved to Milan, by then a centre of medical investigation1. The vast majority of his studies of the heart were produced in Milan from 1508 to 1513, though he continued to work on them later in Rome11.

Working in monastery hospitals and medical schools, he dissected perhaps thirty human corpses, and many more animals, recording the findings in hundreds of detailed drawings and thousands of words10. By his own count he dissected thirty corpses in his lifetime — in Milan, at hospitals in Florence and Rome, and in Pavia1. There he collaborated with the physician-anatomist Marcantonio della Torre1. That collaboration gave his studies a new dimension1. He outlined a plan for a comprehensive work covering exact reproductions of the body, comparative anatomy and physiology, hoping to finish it in the winter of 1510–111. To explore the heart he injected warm wax into a bull's heart to examine the valves' forms, and built a glass model to study the hydraulics of the blood11.

Had he published his researches, his intended treatise would have been the single most important work on anatomy ever written10. Perfectionism, difficulties reconciling observations with traditional beliefs, and bad luck prevented its completion10. During his lifetime the medical investigations remained private — he did not consider himself a professional in anatomy and neither taught nor published110. The drawings and translated notes reached print only in the late nineteenth century110. Today almost the complete set of the anatomical drawings and comments is owned by the British Crown and held in the Royal Library at Windsor Castle11.

In the late 19th century the anatomical drawings were finally published and the notes translated, and we can now understand Leonardo as one of the greatest anatomists ever to have lived.10

Royal Collection Trust — Leonardo's Study of Anatomy

His findings from these studies were recorded in the famous anatomical drawings, which are among the most significant achievements of Renaissance science.1

Encyclopaedia Britannica — Leonardo da Vinci

The Mona Lisa: a documented history

Leonardo began the portrait now known as the Mona Lisa about 1503, without doubt for the Florentine gentleman Francesco del Giocondo12. His wife, Lisa Gherardini (1479–1542), is the sitter named in the Louvre's own catalogue12. The painting was already begun by October 150312. A document in a Heidelberg library, discovered in 2005, preserves a handwritten note by Agostino Vespucci, a collaborator of Machiavelli, mentioning 'the head of Lisa del Giocondo'12. Leonardo kept the portrait with him, its execution still unfinished at his death, and it was most probably acquired by Francis I in 151812.

He painted it on a thin poplar panel of about 79.4 by 53.4 centimetres, to show the sitter at natural scale12. Over the years the wood warped slightly and cracked12. An eleven-centimetre split descends from the top of the panel, crossing the scalp and stopping at the forehead12. The softly sculptural face shows his handling of sfumato, a technique made possible by the relatively new oil paint in Italy13. Part of the royal collection before it became the property of the French people during the Revolution, the painting has belonged to the Louvre's collection since 18041312. The museum's catalogue gives an acquisition date of 17931312.

Mona Lisa by Leonardo da Vinci, oil on poplar panel
Mona Lisa (Portrait of Lisa Gherardini), oil on poplar panel, begun c. 1503; Musée du Louvre, Paris (INV 779). · Leonardo da Vinci (C2RMF photographic reproduction) · Public domain
DateEvent
c. 1503The portrait is begun — without doubt for Francesco del Giocondo.12
October 1503Agostino Vespucci notes 'the head of Lisa del Giocondo'.12
1518Most probably acquired by Francis I.12
1793 / 1804Louvre catalogue acquisition date 1793; in the museum's collection since 1804.1213
21 August 1911Stolen from the Louvre.14
1913Recovered; it becomes the most famous painting in the world.12
1956A vandal's stone breaks the glass; the left elbow is slightly damaged.12
1963Shown in the United States: around 40,000 visitors a day.13
1966Installed in the Salle des États, Room 711, Denon wing.14
2005Placed in a protective glass case.14
A documented timeline of the Mona Lisa's public life.

In 1911 the painting was stolen — by Vincenzo Peruggia, a glazier who had worked at the museum13. The theft caused an immediate media sensation13. People flocked to see the empty space, the museum's director of paintings resigned, and Guillaume Apollinaire and Pablo Picasso were even arrested as suspects13. After its recovery in 1913 it became the Louvre's principal masterpiece and the most famous work in the world12. During the Second World War it was singled out as the most endangered artwork in the Louvre and evacuated to the French countryside, returning in 194513. Theories about the sitter — that she was Leonardo's mother, Caterina, or that the painting is a self-portrait — remain unproven13. Attempts to settle the debate by testing Lisa del Giocondo's DNA were inconclusive13.

Salvator Mundi and the attribution debates

Salvator Mundi (c. 1500) is the panel at the centre of modern attribution argument1. Historians in the 21st century believe Leonardo likely contributed to it, and in 2017 it sold for a record-breaking $450.3 million at auction1. The panel was marketed as a wholly Leonardo work, yet some scholars believe the master painted just a few passages, such as the blessing hand and the tendrils of hair1. The sale overlooked its poor condition, which required extensive conservation1. The episode attested to Leonardo's enduring celebrity — and to his powerful position in the canon of art history centuries after his death1.

The role of most of his studio associates is unclear, which leads to the question of the so-called apocryphal works made with their collaboration1. Scholars have been unable to agree in their attributions1. The documented core is small1. Only seventeen of the paintings that have survived can be definitely attributed to him, and several of them are unfinished1.

Myth-audit: 'universal genius', the helicopter, and 'the first scientist'

The legend has a founding text: Giorgio Vasari's Lives of the Most Eminent Painters, Sculptors, and Architects15. Published in 1550, it became a standard work of art history with its famous attribution of superhuman qualities to Leonardo15. Vasari's sketch of his unique genius helped crystallize an evolving theory of exceptional human ability, later labelled 'universal genius', with da Vinci as its poster child15. Though the label itself, as JSTOR Daily notes, is not a term of precision15. Britannica's own account begins by calling Leonardo the figure whose skill and intelligence, perhaps more than that of any other, epitomized the Renaissance humanist ideal1.

Sometimes, in supernatural fashion… beauty, grace, and talent are united beyond measure in one single person…15

Giorgio Vasari, Lives of the Most Eminent Painters, Sculptors, and Architects (1550)

…skill and intelligence, perhaps more than that of any other figure, epitomized the Renaissance humanist ideal.1

Encyclopaedia Britannica — Leonardo da Vinci

What the record supports is remarkable without exaggeration1. The 'helical airscrew' he designed around 1487 almost seems a prototype for the modern helicopter1. But it lacked an adequate source of power for propulsion and lift — nothing flew1. He dissected perhaps thirty corpses and recorded what he saw in hundreds of drawings110. Yet he neither taught nor published his medical findings in his lifetime110. The anatomical drawings reached print only in the late nineteenth century110. And only seventeen of the surviving paintings can be definitely attributed to him1.

Final years, death, and the surviving record

In 1513 — with the French temporarily expelled from Milan — the sixty-year-old Leonardo moved to Rome1. There his patron Giuliano de' Medici, brother of the new pope Leo X, gave him a suite of rooms in the Belvedere and a considerable monthly stipend1. No large commissions followed1. He worked on mathematical studies, technical experiments and surveys of ancient monuments, and his letters read as embittered1.

Perhaps stifled by the Roman scene, at sixty-five he accepted King Francis I's invitation and left Italy forever at the end of 1516 with Melzi, his most devoted pupil1. He spent his last three years at the small residence of Cloux, later called Clos-Lucé, near Amboise1. He bore the title Premier peintre, architecte et méchanicien du Roi — 'First painter, architect, and engineer to the King'1. The king treated him as an honoured guest, granting him freedom of action1. He died at Cloux on 2 May 1519 and was buried in the palace church of Saint-Florentin, devastated in the French Revolution and torn down in the early nineteenth century1. His grave can no longer be located1. The manuscripts passed to Melzi1. Seventeen attributed paintings survive him; and, as Britannica concludes, Leonardo's influence shows little sign of abating1.

Timeline

15 April 1452

Birth at Anchiano, near Vinci

Leonardo is born on 15 April 1452 at Anchiano near Vinci, in the Republic of Florence, to Ser Piero da Vinci, a Florentine notary, and Caterina; his parents are unmarried at the time of his birth.

c. 1467

Entered Verrocchio's workshop

At about the age of 15 Leonardo's father apprentices him to Andrea del Verrocchio, whose renowned workshop gives him a multifaceted training in painting, sculpture and the technical-mechanical arts; he also works in the next-door workshop of Antonio Pollaiuolo.

1472

Admitted to the painters' guild of Florence

Leonardo is accepted into the painters' guild of Florence; he remains in his teacher's workshop for five more years and then works independently in Florence until 1481.

1482

Moves to Milan in the service of Ludovico Sforza

Leonardo moves to Milan to work in the service of the city's duke, giving up the Adoration of the Magi commission; he is listed as pictor et ingeniarius ducalis ('painter and engineer of the duke') and spends 17 years in Milan, until Ludovico's fall from power in 1499.

c. 1483

Begins The Virgin of the Rocks

He probably begins The Virgin of the Rocks about 1483 as an altarpiece for the church of San Francesco Grande, Milan; the project leads to 10 years of litigation with the Confraternity of the Immaculate Conception and to a second version of the work that he completed in 1508.

c. 1490

First anatomy campaign

In the first of two campaigns of anatomical research — around 1490, and again between 1506 and 1513 — Leonardo investigates the nervous system, the internal organs, the bones and muscles, the heart and the reproductive systems; working in monastery hospitals and medical schools he dissects perhaps thirty human corpses, recording the findings in hundreds of drawings.

1495–98

The Last Supper painted

Commissioned by Ludovico Sforza, the wall painting of the Last Supper is executed probably between 1495 and 1498 for the Dominican monastery Santa Maria delle Grazie in Milan; Leonardo bypasses true fresco and uses an experimental tempera or oil technique on two layers of dry preparatory ground, so the pigments are not permanently attached and the painting begins to flake within a few years.

1503

Begins the Mona Lisa

Leonardo begins the Mona Lisa about 1503, probably for the Florentine gentleman Francesco del Giocondo; an October 1503 marginal note by Agostino Vespucci refers to 'the head of Lisa del Giocondo'; he keeps the portrait with him, still unfinished, until his death and it was probably acquired by Francis I in 1518.

1508–13

Second Milanese period

Back in Milan in the service of the French governors, Leonardo's anatomical studies achieve a new dimension in collaboration with Marcantonio della Torre, a famous anatomist from Pavia, and he plans to finish his anatomical manuscript in the winter of 1510–11.

1513–16

Rome, under Giuliano de' Medici

The now 60-year-old Leonardo moves to Rome, where his patron Giuliano de' Medici gives him a suite of rooms in the Belvedere, in the Vatican, and a considerable monthly stipend; no large commissions follow, and he works on mathematical studies, technical experiments and surveys of ancient monuments.

1516

Leaves Italy for France

At the invitation of King Francis I, Leonardo leaves Italy for the last time at the end of 1516 together with his pupil Melzi; he spends the last three years of his life at the small residence of Cloux (later called Clos-Lucé) near Amboise, bearing the title Premier peintre, architecte et méchanicien du Roi.

2 May 1519

Death at Cloux; Melzi inherits the notebooks

Leonardo dies at Cloux and is buried in the palace church of Saint-Florentin, later torn down, so that his grave can no longer be located; after his death his pupil and friend Melzi becomes heir to more than 10,000 of Leonardo's closely written and illustrated pages.

21 August 1911

The theft of the Mona Lisa

On 21 August 1911 panic broke out at the Louvre as the Mona Lisa disappeared; the glazier Vincenzo Peruggia, who had worked at the museum, later tried to sell the painting to an Italian art dealer, who alerted the authorities; recovered in 1913, the painting's fame grew all the greater.

1999

Last Supper restoration completed

An extensive and controversial 20-year restoration of the Last Supper is completed in 1999; restorers remove previous retouches, grime and varnish, but many critics argue that very little of Leonardo's original work is left; visitors are today allotted 15 minutes to view the mural in small groups.

Frequently asked questions

Where and when was Leonardo da Vinci born?

Leonardo da Vinci was born on 15 April 1452 at Anchiano, near Vinci in the Republic of Florence, and died on 2 May 1519 at Cloux — now Clos-Lucé — in France.1 His parents were unmarried at the time of his birth.1

Who was Leonardo's mother?

She was possibly Caterina di Meo Lippi, a young peasant woman who may have been orphaned at a young age; a few scholars have posited instead that she was an enslaved woman from Asia.1

What training did Leonardo receive, and when did he join the painters' guild?

At about the age of fifteen his father apprenticed him to Andrea del Verrocchio, in whose renowned workshop he received a multifaceted training that included painting and sculpture as well as the technical-mechanical arts.1 In 1472 he was accepted into the painters' guild of Florence, remained with Verrocchio five more years, and then worked independently in Florence until 1481.1

Why has the Last Supper deteriorated so badly?

Leonardo used an experimental technique of tempera or oil paint on two layers of dry preparatory ground, so the pigments were never permanently attached to the wall and the painting began to flake within a few years.4 It then suffered kitchen steam and smoke, candle soot, dampness, a door cut through the wall in 1652, a nineteenth-century flood and mould, and an Allied bomb in the Second World War; a twenty-year restoration was completed in 1999.4

How many dissections did Leonardo perform, and when?

In two campaigns of research — around 1490, and between 1506 and 1513 — working in monastery hospitals and medical schools, he dissected perhaps thirty human corpses.10 By his own count he dissected thirty corpses in his lifetime.1

Where are Leonardo's notebooks and drawings today?

At his death in 1519 his drawings and notebooks were bequeathed to Francesco Melzi, and after Melzi's death around 1570 the collection passed to Pompeo Leoni, who pasted the sheets into at least two large albums.6 One became the Codex Atlanticus, preserved in the Veneranda Biblioteca Ambrosiana in Milan, while the Royal Collection at Windsor holds around 550 of Leonardo's drawings, the most important group in the world.6

When was the Mona Lisa begun, and has the sitter's identity been proven?

The work was already begun by October 1503, as a document in a Heidelberg library discovered in 2005 attests.12 The sitter's identity has not been definitively proven, and attempts in the 21st century to settle the debate by testing Lisa del Giocondo's DNA were inconclusive.13

Did Leonardo invent the helicopter?

He designed a 'helical airscrew' around 1487 that almost seems a prototype for the modern helicopter, but, like the other vehicles he designed, it lacked an adequate source of power to provide propulsion and lift.1

Knowledge graph

apprenticed_toapprenticed_toservedservedpaintedpaintedpaintedpaintedcontributed_tocontributed_tobequeathedbequeathedpassed_topassed_toassembledassembledlocated_inlocated_incommissioned_bycommissioned_byapprenticed_toapprenticed_toservedservedcommissionedcommissionedpaintedpaintedpaintedpaintedcontributed_tocontributed_toinherited_frominherited_fromapprenticed_toapprenticed_toservedservedpaintedpaintedpaintedpaintedcontributed_tocontributed_tobequeathedbequeathedpassed_topassed_toassembledassembledlocated_inlocated_incommissioned_bycommissioned_byapprenticed_toapprenticed_toservedservedcommissionedcommissionedpaintedpaintedpaintedpaintedcontributed_tocontributed_toinherited_frominherited_fromAndrea del VerrocchioAndrea del VerrocchioCodex AtlanticusCodex AtlanticusFrancesco MelziFrancesco MelziFrancesco del GiocondoFrancesco del GiocondoLudovico SforzaLudovico SforzaMona LisaMona LisaPompeo LeoniPompeo LeoniRoyal Library, WindsorRoyal Library, WindsorSalvator MundiSalvator MundiThe Last SupperThe Last SupperVeneranda Biblioteca AmbrosianaVeneranda Biblioteca AmbrosianaLeonardo da VinciLeonardo da Vinci

Sources & citations

Every factual claim in this article is drawn from the sources below. Bracketed numbers in the text link to the corresponding source.

  1. 1
    Leonardo da Vinci | Biography, Art, Paintings, Mona Lisa, Drawings, Inventions, & FactsEncyclopaedia BritannicaReferenceAccessed 2026-09-17© Encyclopaedia Britannica
  2. 2
    The Baptism of Christ by Verrocchio and LeonardoUffizi GalleriesPrimary sourceAccessed 2026-09-17© Uffizi Galleries
  3. 3
    Annunciation by Leonardo da VinciUffizi GalleriesPrimary sourceAccessed 2026-09-17© Uffizi Galleries
  4. 4
    Last Supper | History, Painting, Milan, Leonardo da Vinci, & FactsEncyclopaedia BritannicaReferenceAccessed 2026-09-17© Encyclopaedia Britannica
  5. 5
    Last Supper, Leonardo Da Vinci — Cenacolo Vinciano Official WebsiteMuseo del Cenacolo Vinciano, Milan (Pinacoteca di Brera)Primary sourceAccessed 2026-09-17© Museo del Cenacolo Vinciano
  6. 6
    Leonardo in the Royal Collection (Collection Story)Royal Collection TrustPrimary sourceAccessed 2026-09-17© Royal Collection Enterprises Limited
  7. 7
    leonardo//thek@ — Introduction (Codex Atlanticus digital library)Museo Galileo, FlorencePrimary sourceAccessed 2026-09-17© Museo Galileo — Istituto e Museo di Storia della Scienza
  8. 8
    Water as Microscope of Nature. Leonardo da Vinci's Codex Leicester (exhibition page)Uffizi GalleriesPrimary sourceAccessed 2026-09-17© Uffizi Galleries
  9. 9
    Codex AtlanticusVeneranda Biblioteca AmbrosianaPrimary sourceAccessed 2026-09-17© Veneranda Biblioteca Ambrosiana
  10. 10
    Leonardo's Study of Anatomy (Collection Story)Royal Collection TrustPrimary sourceAccessed 2026-09-17© Royal Collection Enterprises Limited
  11. 11
    Cardiac anatomy in the 'Dreyfus Madonna' by Leonardo da Vinci (Interact Cardiovasc Thorac Surg, 2020)Oxford University Press / PubMed Central (NLM)Primary sourceAccessed 2026-09-17© The Author(s) 2020; published by Oxford University Press on behalf of the European Association for Cardio-Thoracic Surgery
  12. 12
    Portrait de Lisa Gherardini, épouse de Francesco del Giocondo, dit La Joconde ou Monna Lisa (INV 779)Musée du Louvre — Collections databasePrimary sourceAccessed 2026-09-17© Musée du Louvre
  13. 13
    Mona Lisa | Painting, Subject, History, Meaning, & FactsEncyclopaedia BritannicaReferenceAccessed 2026-09-17© Encyclopaedia Britannica
  14. 14
    From the 'Mona Lisa' to 'The Wedding Feast at Cana' — The Salle des ÉtatsMusée du LouvrePrimary sourceAccessed 2026-09-17© Musée du Louvre
  15. 15
    The Destructive Myth of the Universal GeniusJSTOR DailyReferenceAccessed 2026-09-17© JSTOR Daily (ITHAKA)